Meiotic chromosome movements in plants, a puppet show?

Meiosis is a special type of cell division by which sexually reproducing organisms maintain their chromosome number across generations. This process produces haploid gametes by two successive rounds of cell division preceded by a unique DNA replication event. During the first meiotic prophase, the homologous chromosomes form stable bivalents, a process which implies their recognition, with a subsequent step of intimate alignment (pairing), synapsis (physical association of paired chromosomes by the synaptonemal complex, SC), and recombination (exchange of chromosomal regions). Usually, homologous chromosomes are physically separated at initiation of prophase I. In many organisms, telomeres initiate a non-random movement at the entrance of meiosis that brings homologs together tethering at the inner surface of the nuclear envelope (NE). Sad1/UNC-84 (SUN)-domain proteins are inner NE proteins involved in complexes that link cytoskeletal elements